HPV reference clones – International Human Papillomavirus Reference Center

Officialy established HPV types

Virus name	Genus name	Species name	GenBank ID	Date submitted	Submitted by	Reference
HPV1	Mu	Mu-1	V01116	1984	L. Gissman	Danos et al, 1982
HPV2	Alpha	Alpha-4	X55964	1984	L. Gissman	Hirsch-Behnam et al, 1990
HPV3	Alpha	Alpha-2	X74462	1984	L. Gissman	Delius and Hofmann, 1994
HPV4	Gamma	Gamma-1	X70827	1984	L. Gissman	Egawa et al, 1993
HPV5	Beta	Beta-1	M17463	1984	L. Gissman	Zachow et al, 1987
HPV6	Alpha	Alpha-10	X00203	1984	L. Gissman	Schwarz et al, 1983
HPV7	Alpha	Alpha-8	X74463	1984	L. Gissman and T. Oltersdorf	Delius and Hofmann, 1994
HPV8	Beta	Beta-1	M12737	1984	L. Gissman	Fuchs et al, 1986
HPV9	Beta	Beta-2	X74464	1984	L. Gissman	Delius and Hofmann, 1994
HPV10	Alpha	Alpha-2	X74465	1984	L. Gissman	Delius and Hofmann, 1994
HPV11	Alpha	Alpha-10	M14119	1984	L. Gissman	Dartmann et al, 1986
HPV12	Beta	Beta-1	X74466	1984	G. Orth via L. Gissman	Delius and Hofmann, 1994
HPV13	Alpha	Alpha-10	X62843	1984	L. Gissman	Van Ranst et al, 1992
HPV14	Beta	Beta-1	X74467	1984-10-22	G. Orth	Delius and Hofmann, 1994
HPV15	Beta	Beta-2	X74468	1984-10-09	G. Orth	Delius and Hofmann, 1994
HPV16	Alpha	Alpha-9	K02718	1984	M. Durst and L. Gissman	Seedorf et al, 1986
HPV17	Beta	Beta-2	X74469	1984-10	G. Orth	Delius and Hofmann, 1994
HPV18	Alpha	Alpha-7	X05015	1984	M. Boshart	Cole and anos, 1987
HPV19	Beta	Beta-1	X74470	1984-10	G. Orth	Delius and Hofmann, 1994
HPV20	Beta	Beta-1	U31778	1990-05-18	G. Orth	Kremsdorf et al, 1984
HPV21	Beta	Beta-1	U31779	1984-10	G. Orth	Kremsdorf et al, 1984
HPV22	Beta	Beta-2	U31780	1984	G. Orth	Kremsdorf et al, 1984
HPV23	Beta	Beta-2	U31781	1984-10	G. Orth	Kremsdorf et al, 1984
HPV24	Beta	Beta-1	U31782	1984-10	G. Orth	Kremsdorf et al, 1984
HPV25	Beta	Beta-1	X74471	1984-09-19	L. Gissman and H. Pfister	Delius and Hofmann, 1994
HPV26	Alpha	Alpha-5	X74472	1985-04-10	R. Ostrow	Delius and Hofmann, 1994
HPV27	Alpha	Alpha-4	X74473		R. Ostrow	Delius and Hofmann, 1994
HPV28	Alpha	Alpha-2	U31783	1986-04-23	G. Orth	Delius,H. Unpublished
HPV29	Alpha	Alpha-2	U31784	1986-04	G. Orth	Delius,H. Unpublished
HPV30	Alpha	Alpha-6	X74474	1985-04		Delius and Hofmann, 1994
HPV31	Alpha	Alpha-9	J04353	1985-02	A. Lorincz	Goldsborough at al, 1989
HPV32	Alpha	Alpha-1	X74475	1986-04	G. Orth	Delius and Hofmann, 1994
HPV33	Alpha	Alpha-9	M12732	1985-12-85	G. Orth	Cole and Streeck, 1986
HPV34	Alpha	Alpha-11	X74476	1985-12-03	G. Orth	Delius and Hofmann, 1994
HPV35	Alpha	Alpha-9	X74477	1986-10	A. Lorincz	Delius and Hofmann, 1994
HPV36	Beta	Beta-1	U31785	1986	G. Orth	Kawashima et al, 1986
HPV37	Beta	Beta-2	U31786	1986	W. Scheurlen	Scheurlen et al, 1986
HPV38	Beta	Beta-2	U31787	1986	W. Scheurlen	Scheurlen et al, 1986
HPV39	Alpha	Alpha-7	M62849	1987-09-21	G. Orth	Volpers and Streeck, 1991
HPV40	Alpha	Alpha-8	X74478	1987-09-21	E.M. de Villiers	Delius and Hofmann, 1994
HPV41	Nu	Nu-1	X56147	1987-09-21	E.M. de Villiers	Hirt et al, 1991
HPV42	Alpha	Alpha-1	M73236	1987-09-21	G. Orth	Philipp et al, 1992
HPV43	Alpha	Alpha-8	AJ620205	1987-07-29	A. Lorincz	Lorincz et al, 1989
HPV44	Alpha	Alpha-10	U31788	1987-07-29	A. Lorincz	Delius,H., Unpublished
HPV45	Alpha	Alpha-7	X74479	1986-03-28	Z. Naghashfar	Delius and Hofmann, 1994
"HPV46"			re-classified as HPV20 subtype
HPV47	Beta	Beta-1	M32305	1987-03-27	M. Ishibashi	Kiyono et al, 1990
HPV48	Gamma	Gamma-2	U31789	1987-11-17	L. Gissman	Muller et al, 1989
HPV49	Beta	Beta-3	X74480	1987-04-21	M. Favre	Delius and Hofmann, 1994
HPV50	Gamma	Gamma-3	U31790	1987-09-21	M. Favre	Favre et al, 1989
HPV51	Alpha	Alpha-5	M62877	1987-01-06	C. Crum	Lungu et al, 1991
HPV52	Alpha	Alpha-9	X74481	1987-05	W. Lancaster	Delius and Hofmann, 1994
HPV53	Alpha	Alpha-6	X74482	1987-11-17	L. Gissman	Delius and Hofmann, 1994
HPV54	Alpha	Alpha-13	U37488	1987-11-10	G. Orth	Delius,H., Unpublished
"HPV55"			re classified as HPV44 subtype	1987-11-10	M. Favre
HPV56	Alpha	Alpha-6	X74483	1987-09-30	A. Lorincz	Delius and Hofmann, 1994
HPV57	Alpha	Alpha-4	X55965	1989-06-26	E.M. de Villiers	Hirsch-Behnam et al, 1990
HPV58	Alpha	Alpha-9	D90400	1988-06-26	T. Matsukura	Kirii et al, 1991
HPV59	Alpha	Alpha-7	X77858	1989-01-02	T. Matsukura	Rho et al, 1994
HPV60	Gamma	Gamma-4	U31792	1989-01-02	T. Matsukura	Matsukura et al, 1992
HPV61	Alpha	Alpha-3	U31793	1989-03-01	T. Matsukura	Delius,H. Unpublished
HPV62	Alpha	Alpha-3	AY395706	1998-03-01	T. Matsukura	Fu et al, 2004
HPV63	Mu	Mu-2	X70828	1991-04-11	K. Egawa	Egawa et al, 1993
"HPV64"			re-classified as HPV34 subtype
HPV65	Gamma	Gamma-1	X70829	1989-01-02	T. Matsukura	Egawa et al, 1993
HPV66	Alpha	Alpha-6	U31794	1981-08-07	G. Orth	Delius,H., Unpublished
HPV67	Alpha	Alpha-9	D21208	1989-10-30	T. Matsukura	Kirii et al, 1998
HPV68	Alpha	Alpha-7	X67161	1981-03-14	G. Orth	Longuet et al, 1996
HPV69	Alpha	Alpha-5	AB027020	1991-09-09	T. Matsukura	Kino et al, 2000
HPV70	Alpha	Alpha-7	U21941	1993-02-19	G. Orth	Forslund and Hansson, 1996
HPV71	Alpha	Alpha-14	AB040456	1991-09-09	T. Matsukura	Matsukura and Sugase, 2001
HPV72	Alpha	Alpha-3	X94164	1993-10-11	Y. He	Volter et al, 1996
HPV73	Alpha	Alpha-11	X94165	1993-10-11	Y. He	Volter et al, 1996
HPV74	Alpha	Alpha-10	AF436130	1993-09-27	G. Orth	Burk and Terai, Unpublished
HPV75	Beta	Beta-3	Y15173	1994-10-05	V. Shamanin and E.M. de Villiers	Delius et al, 1998
HPV76	Beta	Beta-3	Y15174	1994-10-05	E.M. de Villiers	Delius et al, 1998
HPV77	Alpha	Alpha-2	Y15175	1995-07-31	V. Shamanin	Delius et al, 1998
HPV78	Alpha	Alpha-2	AB793779	1995-01-03	T. Matsukura	Sata et al, unpublished
"HPV79"			replaced by HPV91	1996-12-30	G. Orth	Terai and Burk, 2002
HPV80	Beta	Beta-2	Y15176	1995-07-31	K. Bergmann	Delius et al, 1998
HPV81	Alpha	Alpha-3	AJ620209	1996-08-06	T. Matsukura	Matsukura and Sugase, 2001
HPV82	Alpha	Alpha-5	AB027021	1997-01-20	T. Matsukura	Kino et al, 2000
HPV83	Alpha	Alpha-3	AF151983	1998-06-12	K.H. Fife and D. Brown	Brown et al, 1999
HPV84	Alpha	Alpha-3	AF293960	2000-04-05	R. Burk	Terai and Burk, 2001
HPV85	Alpha	Alpha-7	AF131950	1999-09-02	V. Chow	Chow and Leong, 1999
HPV86	Alpha	Alpha-3	AF349909	2000-09-01	R. Burk	Terai and Burk, 2001
HPV87	Alpha	Alpha-3	AJ400628	2000-08-31	S. Menzo	Menzo et al, 2001
HPV88	Gamma	Gamma-5	EF467176	2001-07-30	K. Egawa	Kullander et al, 2008
HPV89	Alpha	Alpha-3	AF436128	2001-08-27	R. Burk	Terai and Burk, 2002
HPV90	Alpha	Alpha-14	AY057438	2001-08-27	R. Burk	Terai and Burk, 2002
HPV91	Alpha	Alpha-8	AF419318	2001-08-27	R. Burk	Terai and Burk, 2002
HPV92	Beta	Beta-4	AF531420	2001-10-12	O. Forslund	Forslund et al, 2003
HPV93	Beta	Beta-1	AY382778	2002-02-12	O. Forslund	Vasiljevic et al, 2007
HPV94	Alpha	Alpha-2	AJ620211	2002-10-23	E.M. de Villiers	de Villiers et al, 2009
HPV95	Gamma	Gamma-1	AJ620210	2003-06-05	E.M. de Villiers and K. Egawa	Egawa et al, 2005
HPV96	Beta	Beta-5	AY382779	2002-10-04	O. Forslund	Vasiljevic et al, 2007
HPV97	Alpha	Alpha-7	DQ080080	2004-07-21	R. Burk	Chen and Burk, 2007
HPV98	Beta	Beta-1	FM955837	2004-07-15	E.M. de Villiers	de Villiers and Gunst, 2009
HPV99	Beta	Beta-1	FM955838	2004-07-15	E.M. de Villiers	de Villiers and Gunst, 2009
HPV100	Beta	Beta-2	FM955839	2004-07-26	E.M. de Villiers	de Villiers and Gunst, 2009
HPV101	Gamma	Gamma-6	DQ080081	2004-07-21	R. Burk	Chen et al, 2007
HPV102	Alpha	Alpha-3	DQ080083	2004-09-15	R. Burk	Narechania et al, 2005
HPV103	Gamma	Gamma-6	DQ080078	2004-09-15	R. Burk	Chen et al, 2007
HPV104	Beta	Beta-2	FM955840	2004-04-15	E.M de Villiers	de Villiers and Gunst, 2009
HPV105	Beta	Beta-1	FM955841	2004-08-04	E.M. de Villiers	de Villiers and Gunst, 2009
HPV106	Alpha	Alpha-14	DQ080082	2004-07-21	R. Burk	Narechania et al, 2005
HPV107	Beta	Beta-2	EF422221	2006-11-03	J. Kullander and O. Forslund	Vasiljevic et al, 2008
HPV108	Gamma	Gamma-6	FM212639	2006-11	R. Nobre and E.M. de Villiers	Nobre et al, 2009
HPV109	Gamma	Gamma-7	EU541441	2007-08-23	J. Kullander and O. Forslund	Ekstrom et al, 2010
HPV110	Beta	Beta-2	EU410348	2007-09-28	N. Vasiljevic and O. Forslund	Vasiljevic et al, 2008
HPV111	Beta	Beta-2	EU410349	2007-11-05	N. Vasiljevic and O. Forslund	Vasiljevic et al, 2008
HPV112	Gamma	Gamma-8	EU541442	2007-12-21	J. Kullander and O. Forslund	Ekstrom et al, 2010
HPV113	Beta	Beta-2	FM955842	2008-03-25	E.M. de Villiers	de Villiers and Gunst, 2009
HPV114	Alpha	Alpha-3	GQ244463	2008-11-14	J. Kullander and O. Forslund	Ekstrom et al, 2010
HPV115	Beta	Beta-3	FJ947080	2008-12-08	A. Giri	Chouhy et al, 2010
HPV116	Gamma	Gamma-9	FJ804072	2009-01-13	L. Li and E. Delwart	Li et al, 2009
HPV117	Alpha	Alpha-2	GQ246950	2009-07-02	I. Nindl	Kohler et al, 2010
HPV118	Beta	Beta-1	GQ246951	2009-07-02	I. Nindl	Kohler et al, 2010
HPV119	Gamma	Gamma-8	GQ845441	2009-07-15	R. Burk	Bernard et al, 2010
HPV120	Beta	Beta-2	GQ845442	2009-07-15	R. Burk	Bernard et al, 2010
HPV121	Gamma	Gamma-10	GQ845443	2009-07-15	R. Burk	Bernard et al, 2010
HPV122	Beta	Beta-2	GQ845444	2009-07-15	R. Burk	Bernard et al, 2010
HPV123	Gamma	Gamma-7	GQ845445	2009-07-15	R. Burk	Bernard et al, 2010
HPV124	Beta	Beta-1	GQ845446	2009-07-15	R. Burk	Bernard et al, 2010
HPV125	Alpha	Alpha-2	FN547152	2009-09-08	A. Kovanda, B. Kocjan and M. Poljak	Kovanda et al, 2011
HPV126	Gamma	Gamma-11	AB646346	2010-09-09	N. Egawa	Egawa et al, 2012
HPV127	Gamma	Gamma-12	HM011570	2009-11-20	C. Buck	Schowalter et al, 2010
HPV128	Gamma	Gamma-13	GU225708	2009-12-02	I. Nindl	Kohler et al, 2011
HPV129	Gamma	Gamma-9	GU233853	2009-12-02	I. Nindl	Kohler et al, 2011
HPV130	Gamma	Gamma-10	GU117630	2009-12-02	I. Nindl	Kohler et al, 2011
HPV131	Gamma	Gamma-14	GU117631	2009-12-02	I. Nidl	Kohler et al, 2011
HPV132	Gamma	Gamma-12	GU117632	2009-12-02	I. Nidl	Kohler et al, 2011
HPV133	Gamma	Gamma-10	GU117633	2009-12-02	I. Nidl	Kohler et al, 2011
HPV134	Gamma	Gamma-7	GU117634	2009-12-02	I. Nidl	Kohler et al, 2011
HPV135	Gamma	Gamma-15	HM999987	2009-12-07	R. Burk	Bottalico et al, 2011
HPV136	Gamma	Gamma-11	HM999988	2009-12-07	R. Burk	Bottalico et al, 2011
HPV137	Gamma	Gamma-16	HM999989	2009-12-07	R. Burk	Bottalico et al, 2011
HPV138	Gamma	Gamma-7	HM999990	2009-12-07	R. Burk	Chen et al, Unpublished
HPV139	Gamma	Gamma-7	HM999991	2009-12-07	R. Burk	Chen et al, Unpublished
HPV140	Gamma	Gamma-11	HM999992	2009-12-23	R. Burk and Z. Chen	Bottalico et al, 2011
HPV141	Gamma	Gamma-11	HM999993	2009-12-23	R. Burk and Z. Chen	Bottalico et al, 2011
HPV142	Gamma	Gamma-10	HM999994	2009-12-23	R. Burk and Z. Chen	Chen et al, Unpublished
HPV143	Beta	Beta-1	HM999995	2010-02-15	R. Burk and Z. Chen	Bottalico et al, 2011
HPV144	Gamma	Gamma-17	HM999996	2010-02-15	R. Burk and Z. Chen	Bottalico et al, 2011
HPV145	Beta	Beta-2	HM999997	2010-02-15	R. Burk and Z. Chen	Bottalico et al, 2011
HPV146	Gamma	Gamma-15	HM999998	2010-02-15	R. Burk and Z. Chen	Chen et al, Unpublished
HPV147	Gamma	Gamma-8	HM999999	2010-02-15	R. Burk and Z. Chen	Chen et al, Unpublished
HPV148	Gamma	Gamma-12	GU129016	2009-12-02	I. Nidl	Kohler et al, 2011
HPV149	Gamma	Gamma-7	GU117629	2009-12-02	I. Nidl	Kohler et al, 2011
HPV150	Beta	Beta-5	FN677755	2010-02-25	A. Kovanda, B. Kocjan and M. Poljak	Kovanda et al, 2011
HPV151	Beta	Beta-2	FN677756	2010-02-25	A. Kovanda, B. Kocjan and M. Poljak	Kovanda et al, 2011
HPV152	Beta	Beta-1	JF304768	2010-10-14	R. Burk and Z. Chen	Chen et al, unpublished
HPV153	Gamma	Gamma-13	JN171845	2011-02-24	H. Johansson and O. Forslund	Sturegard et al, 2013
HPV154	Gamma	Gamma-11	NC_021483	2011-03-31	O. Forslund and A. Ure	Ure and Forslund, 2014
HPV155	Gamma	Gamma-7	JF906559	2011-05-16	H. Johansson and O. Forslund	Bzhalava et al, 2013
HPV156	Gamma	Gamma-18	JX429973	2011-04-12	D. Chouhy	Chouhy et al, 2013
HPV157	Gamma	Gamma-12	KT698166	2012-05-29	D. Chouhy	Bolatti et al, 2016
HPV158	Gamma	Gamma-1	KT698168	2012-05-29	D. Chouhy	Bolatti et al, 2016
HPV159	Beta	Beta-2	HE963025	2012-07-12	B. Kocjan and M. Poljak	Kocjan et al, 2013
HPV160	Alpha	Alpha-2	AB745694	2012-09-06	T. Kiyono	Mitsuishi et al, 2013
HPV161	Gamma	Gamma-19	JX413109	2012-05-07	J. Li	Li et al, 2012
HPV162	Gamma	Gamma-19	JX413108	2012-05-07	J. Li	Li et al, 2012
HPV163	Gamma	Gamma-20	JX413107	2012-05-07	J. Li	Li et al, 2012
HPV164	Gamma	Gamma-8	JX413106	2012-05-07	J. Li	Li et al, 2012
HPV165	Gamma	Gamma-12	JX444072	2012-05-07	J. Li	Li et al, 2012
HPV166	Gamma	Gamma-19	JX413104	2012-05-07	J. Li	Li et al, 2012
HPV167	Gamma	Gamma-21	NC_022892	2012-05-07	J. Li	Li et al, 2012
HPV168	Gamma	Gamma-8	KC862317	2012-05-07	J. Li	Li et al, 2012
HPV169	Gamma	Gamma-11	JX413105	2012-05-07	J. Li	Li et al, 2012
HPV170	Gamma	Gamma-7	JX413110	2012-05-07	J. Li	Li et al, 2012
HPV171	Gamma	Gamma-11	KF006398	2012-12-08	Q. Feng	Martin et al, 2013
HPV172	Gamma	Gamma-22	KF006399	2012-12-08	Q. Feng	Martin et al, 2013
HPV173	Gamma	Gamma-1	KF006400	2012-12-08	Q. Feng	Martin et al, 2013
HPV174	Beta	Beta-2	HF930491	2013-03-06	B. Kocjan and M. Poljak	Kocjan et al, 2013
HPV175	Gamma	Gamma-23	KC108721	2013-06-18	H. Johansson and O. Forslund	Johansson and Forslund. 2014
HPV176	Gamma	Gamma-8	KR816167	2013-07-02	R. Burk	Chen and Burk. Unpublished.
HPV177	Alpha	Alpha-11	KR816168	2013-07-10	R. Burk	Chen and Burk. Unpublished.
HPV178	Gamma	Gamma-24	KJ130020	2013-07-12	H. Johansson and O. Forslund	Johansson and Forslund. 2014
HPV179	Gamma	Gamma-15	HG421739	2013-07-19	Hosnjak L., B. Kocjan and M. Poljak	Hosnjak et al, 2015
HPV180	Gamma	Gamma-10	KC108722	2013-07-25	H. Johansson and O. Forslund	Johansson and Forslund. 2014
HPV181	Gamma	Gamma-11	KR816169	2013-08-21	R. Burk	Chen and Burk. Unpublished.
HPV182	Beta	Beta-2	KR816170	2013-08-21	R. Burk	Chen and Burk. Unpublished.
HPV183	Gamma	Gamma-20	KR816171	2013-08-21	R. Burk	Chen and Burk. Unpublished.
HPV184	Gamma	Gamma-25	HG530535	2013-09-15	Hosnjak L., B. Kocjan and M. Poljak	Hosnjak et al, 2015
HPV185	Beta	Beta-5	KR816172	2013-10-17	R. Burk	Chen and Burk. Unpublished.
HPV186	Gamma	Gamma-7	KR816173	2013-10-17	R. Burk	Chen and Burk. Unpublished.
HPV187	Gamma	Gamma-26	KR816174	2013-10-17	R. Burk	Chen and Burk. Unpublished.
HPV188	Gamma	Gamma-3	KR816175	2013-10-17	R. Burk	Chen and Burk. Unpublished.
HPV189	Gamma	Gamma-7	KR816176	2013-10-17	R. Burk	Chen and Burk. Unpublished.
HPV190	Gamma	Gamma-24^a	KR816177	2013-10-17	R. Burk	Chen and Burk. Unpublished.
HPV191	Gamma	Gamma-10	KR816178	2013-10-17	R. Burk	Chen and Burk. Unpublished.
HPV192	Gamma	Gamma-15	KR816179	2013-10-17	R. Burk	Chen and Burk. Unpublished.
HPV193	Gamma	Gamma-7	KR816180	2014-01-22	R. Burk	Chen and Burk. Unpublished.
HPV194	Gamma	Gamma-20	KR816181	2014-01-22	R. Burk	Chen and Burk. Unpublished.
HPV195	Beta	Beta-1	KR816182	2014-01-22	R. Burk	Chen and Burk. Unpublished.
HPV196	Beta	Beta-1	KR816183	2014-01-22	R. Burk	Chen and Burk. Unpublished.
HPV197	Gamma	Gamma-24	KM085343	2014-04-14	S. Arroyo	Arroyo et al, 2014
HPV198	Beta	Beta-2	MG921179	2014-04-25	S. Arroyo	Arroyo et al, Unpublished
HPV199	Gamma	Gamma-12	KJ913662	2014-05-19	A. Ostrbenk, B. Kocjan and M. Poljak	Ostrbenk et al, 2015
HPV200	Gamma	Gamma-2	KP692114	2014-08-04	S. Arroyo	Arroyo et al, 2015
HPV201	Gamma	Gamma-27	KP692115	2014-08-04	S. Arroyo	Arroyo et al, 2015
HPV202	Gamma	Gamma-11	KP692116	2014-08-04	S. Arroyo	Arroyo et al, 2015
HPV203	Gamma	Gamma-7	MG921180	2014-10-13	S. Arroyo	Arroyo et al, Unpublished
HPV204	Mu	Mu-3	KP769769	2014-10-22	B. Kocjan and M. Poljak	Kocjan et al, 2015
HPV205	Gamma	Gamma-1	KT698167	2014-12-15	E. Bolatti	Bolatti et al, 2016
HPV206	Beta	Beta-1	U85660	2014-12-22	H. Pfister	Hopfl et al, 1997
HPV207	Gamma	Gamma-15	MK645900	2015-06-25	Bueno
HPV208	Gamma	Gamma-24^a	MK645901	2015-06-25	Bueno
HPV209	Beta	Beta-2	KY242583	2016-02-19	E. Bolatti	Bolatti et al, 2017
HPV210	Gamma	Gamma-12	MH460956	2016-02-19	E. Bolatti	Bolatti et al, 2018
HPV211	Gamma	Gamma-8	MF509816	2016-06-19	T. Meiring	Murahwa et al, 2019
HPV212	Gamma	Gamma-17	MF509817	2016-06-19	T. Meiring	Murahwa et al, 2019
HPV213	Gamma	Gamma-13	MF509818	2016-06-19	T. Meiring	Murahwa et al, 2019
HPV214	Gamma	Gamma-6	MF509819	2016-06-19	T. Meiring	Murahwa et al, 2019
HPV215	Gamma	Gamma-9	MF509820	2016-06-19	T. Meiring	Murahwa et al, 2019
HPV216	Gamma	Gamma-9	MF509821	2016-06-19	T. Meiring	Murahwa et al, 2019
HPV217		re-classified as HPV182 subtype	KY349817	2016-10-02	S. Dutta	Dutta et al, 2017
HPV218		re-classified as HPV189 subtype	KY780961	2016-10-02	S. Dutta	Dutta et al, 2017
HPV219	Gamma	Gamma-13	MH172376	2017-10-10	T. Meiring	Murahwa et al, 2018
HPV220	Gamma	Gamma-17	MH172377	2017-10-10	T. Meiring	Murahwa et al, 2018
HPV221	Gamma	Gamma-10	MH172378	2017-10-10	T. Meiring	Murahwa et al, 2018
HPV222	Gamma	Gamma-19	MH172379	2017-10-10	T. Meiring	Murahwa et al, 2018
HPV223	Gamma	Gamma-22	MG063749	2017-10-16	S. Dutta	Dutta et al, Unpublished
HPV224	Gamma	Gamma-8	MF356498	2017-10-16	S. Dutta	Brancaccio et al, 2017
HPV225	Gamma	Gamma-7	MG520499	2017-10-16	S. Dutta	Dutta et al, Unpublished
HPV226	Gamma	Gamma-6	MG813996	2018-01-24	J. Mossong	Latsuzbaia et al, 2018
HPV227	Beta	Beta-2	MK080568	2018-10-25	R. Brancaccio	Brancaccio et al, 2019
HPV228	Gamma	Gamma-27	ON482334	2019-08-21	C. Boukadida
HPV229	Gamma	Gamma-7	MW535770	2021-09-08	M. Rajeevan
HPV 230	Gamma	Gamma-15	OQ915151	2023-06-23	M. Rajeevan
HPV 231	Gamma	Gamma-10	OP577477	2023-06-23	M. Rajeevan

^a. Assignments are tentative and not official, but might be recommendations to the papillomavirus working group and ICTV.
Last modified by: Laila Sara Arroyo, 2023-09-13
email: sara.arroyo.muhr@ki.se

HPV L1 sequences

A file with all L1 sequences from the officially established HPV types can be found here.

Phylogenetic tree

An updated version of a phylogenetic tree comprising all official HPV types can be downloaded here. Alpha, beta, gamma, mu and nu papillomaviruses are presented in green, blue, red, yellow and purple colors, respectively. The phylogenetic tree is based on the L1 part of the genome. The evolutionary history was inferred by using the Maximum Likelihood method based on the Tamura-Nei model (Tamura and Nei, 1993) and analyses were conducted in MEGA7 (Kumar et al., 2016).